Teaching

My courses emphasize the process of scientific investigation, and whenever possible make connections to environmental issues.

I teach an introductory course for Biology and Environmental Studies majors (Bio 141, Investigations in Ecology and Environmental Science), as well as upper-division courses in ecology (Bio 335, Ecology) and in evolution (Bio 390, Evolution).

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my teaching philosophy

Biology 141

Biology 335

Biology 390

I'm a population biologist, and I work in the areas of ecology, evolution, and conservat ion biology. I teach and advise students in both Biology and Environmental Studies. I have undergraduate degrees in botany and in zoology from the University of Washington in Seattle, and a Ph.D. in ecology and evolutionary biology from Cornell University. I joined the faculty of Lewis and Clark College in 1995, after 15 years at Pomona College. For more information about my professional background, see my vita.

Research

I study the ecology and evolution of wild plant species growing in their natural environments. I'm interested in how environmental factors affect their performance and the evolution of particular traits, especially reproductive strategies.

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Arisaema triphyllum a sex-changing plant

Antennaria spp. plants that reproduce asexually

Linanthus parryae a tiny desert plant with dimorphic flower color

Viola adunca food plant of the threatened Oregon silverspot butterfly