| Dept of Biology, Lewis and Clark College | Dr Kenneth Clifton |
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Biology
221 Lecture Outline |
Lecture 3: Water!
Water's unique properties play an important role in shaping life in the sea.
Seawater is a complex solution with important properties as a solvent, heat sink, and absorber of light.
Water is also unique because its density as a solid is less that as a fluid (ice floats): Why is this important?Water motion in the form of currents, waves, and tides also plays important biological roles.
Water's molecular polarity allows binding with ionsHydrophobic reactions also play important biological roles (e.g. lipid membranes, protein folding, etc.)
"Salinity" reflects a to the concentration of a collection of many inorganic solids within water.The long residency of many elements (Na, Cl, Sr, etc.) keeps them in relatively constant ratio.Major elements (0.1 - 20 ppt): Cl-; Na+; SO4-2; Mg+2; Ca+2; K+; HCO3-
Minor elements (0.1 - .001 ppt): Br-; H2BO3- ; Sr+2; F-
Trace elements (e.g., N, P, Fe) may be biologically limiting
Salinity is influenced by evaporation and dilution
Oxygen dissolves at the sea surface. This, plus photosynthesis, minus respiration, determines local O2 levels
Oxygen dissolves more in cold water, and less in salty water (i.e. lots of O2 at the poles)Anoxic conditions may persist in deep, dark places
It takes 1 calorie to raise 1 gram of water 1 °CTranslation: It takes lots of heat to change water's temperatureWater has a high heat of vaporization
Translation: water absorbs lots of heat before evaporatingBottom line: Water's ability to hold and transfer heat contributes directly to the world's climate
Sunlight hits the earth from different angles and is either reflected or absorbedAbsorption of different wavelengths occurs at different rates
Infrared and ultraviolet attenuate rapidly. Blue penetrates better than redLight intensity diminishes exponentially with depth
Important for photosynthesis and for visual predators seeking preyIntense light levels in shallow water can be bad
Diffusion within seawater allows molecules (nutrients, gasses) to disperse
All else being equal, this should promote uniformityVarious factors may counter the effects of diffusionDifferences in water density (pycnoclines) may inhibit mixing
Temperature plays a role, as warmer, less dense water tends to rise.Solar heat creates thermoclines that vertically stratify water massesSalinity also affects density
Cold, saline waters sinkPressure increases with depth, this changes the solubility of gasses
Water motion may also disrupt or override diffusion
Ways that water moves: currents, tides, wavesCurrents are directional flowsWater movements directly influence patterns of temperature and nutrient availabilityWinds and the Earth's west to east rotation cause global currents
Coriolis effects within fluids: particle deflection on a rotating sphereTo the right in the north and to the left in the south
Westerlies and trade winds are driven by atmospheric pressure differentials
These winds, plus Coriolis effects, create hemispheric gyres of water motion
Clockwise in the north, counterclockwise in the southTry the internet for additional explanations of how coriolis effects influence water motion
Yet another water page, including a bit of animation about coriolis effects
Wind on water along western shores produces upwelling of deeper water
These cold, nutrient-rich waters increase biological productivityCounter currents may occur when water piles up
Strong current reversals (El Nino events) shut down upwellingLarge storms can also cause water to pile up (storm surges)
Not all currents are obvious from the surfaceDeep water currents originate at the poles.They reflect differences in temperature and salinity,
These currents deliver oxygen rich waters to the depths
Back to the top Waves: wind driven "packets" of energy
Why are waves important?Waves are a "predictable" source of disturbance along coastlinesWave energy determines grain size
Most waves originate from wind
Wave size/height (crest to trough) depends on fetch, wind speed, and duration
Other important components of wave energyWavelength (L)Period (T)
Velocity (V)
These are related: V = L/T
Waves break after "feeling bottom"Shoreline topography influences wave energyRefraction may focus energy on headlands
Rip currents sweep away from shore
Longshore currents move down the coast
Seasonal differences in wave energy can shift local topography (berms and bars)
Tsunamis ("tidal waves"): seismic waves have nothing to do with tides
High speed (600 km/hr) = lots of energy!Unpredictable catastrophic events on some coasts causes intermittent temporal disturbance on a massive scale
This video gives a sense of how tsunamis come ashore
Tides generate locally predictable patterns of depth and currentThus, tides influence many aspects of marine biology. For example: The intertidal
Reproduction is also often linked to the tidesWhat causes tides?
Gravity waves driven by the motion of sun and moonRelative celestial position influences tides
Here are two good sites that talk about tides and their predictions:
http://co-ops.nos.noaa.gov/tp4days_old.htmlhttp://www.opsd.nos.noaa.gov/about2.html#ABOUT
Neap tides and Spring tidesPatterns of ebb and flood are also influenced by local topography
In combination, the various properties of water create or organize many different ecological patterns within the sea.